Curriculum Vitae

Dept of Biological Anth and Anatomy Duke University Box 90383 One Science Drive Durham North Carolina 27708

919-660-7328 (office) (919) 382-2969 (home) (email)

• Ph.D., University of California, Davis, CA, 1982


• M.A., University of Minnesota, Minneapolis, MN, 1971


• B.S., Mills College, Oakland, CA, 1969

Areas of Experience

Teaching Summary
Human Origins (Univ. MN, Tulane, CWU, UNO)
Conflict and Conciliation (Freshman writing) (Tulane)
Primate Behavior and Ecology (Tulane, CWU)
Primate Evolution and Adaptation (CWU)
Human Sociobiology (Tulane, CWU, Duke)
Observational Research Methods (Tulane)
Rank (Four field Senior Seminar) (Tulane)
Human Evolution (Tulane, CWU)
Human Ecology (Rollins College)
Cultural Ecology (Rollins College)
Research Methods and Statistics (Rollins College)
Introduction to Cultural Anthropology (Univ. Minnesota)
Human Variation and Adaptation (CWU)
Comparative Animal Behavior
Evolution of Behavior

Professional Experience / Employment History

• Univ. of New Orleans
  Instructor of Anthropology, September, 2003 - June, 2004


• University of New Orleans
  Research Affiliate, Department of Anthropology, September, 2003 - present


• Tulane University
  Faculty of the Graduate School, 1999 - 2002


• Faculty of the Roger Thayer Stone Center for Latin American Studies, 1997 - 2002


• Assistant Professor, July, 1996 - June, 2002


• Rollins College
  Adjunct Professor, Department of Environmental Studies and Department of Anthropology and Sociology, February, 1983 - June, 1984
  Taught undergraduate courses in Human Ecology, Cultural Ecology, Human Sociobiology, Research Design and Statistics


• University of California
  Teaching Assistant, Department of Anthropology, October, 1980 - December, 1981
  Conducted discussion sections, prepared and graded exams, term papers, for Introductory Physical Anthropology. Carried out analysis of data from field research, and completed dissertation, SOCIALIZATION, INFANT MORTALITY, AND INFANT-NONMOTHER INTERACTIONS IN HOWLING MONKEYS (Alouatta palliata) IN COSTA RICA


• Teaching Assistant, Department of Anthropology, September, 1976 - June, 1978
  Conducted discussion sections in Introductory Physical Anthropology, lectured on primate behavior, hominid evolution


• College of Alameda
  Instructor in Anthropology, September, 1973 - July, 1976
  Taught Introduction to Physical Anthropology (Primate Behavior, Hominid Evolution, Population Biology, and Human Variation)


• University of Minnesota
  Teaching Fellow, Department of Anthropology, September, 1969 - June, 1971
  Conducted discussion sections for Cultural and Social Anthropology, Physical Anthropology, and other upper division courses. Edited manuscripts.

Visiting Positions

Visiting Assistant Professor, Duke University, August, 2004 - present

Assistant Professor, Central Washington University, September, 2002 - August, 2003

Awards, Honors, and Distinctions

• Fellow, Newcomb College, Tulane University, 1996-2002


• Who’s Who Among America’s Teachers, 2002


• Barbara Greenbaum Newcomb Fellows Faculty Research Award, 2001-2002


• Mortar Board of Newcomb College Teaching Award, 1999-2000


• Phi Kappa Phi, University of California, Davis, 1980


• Phi Beta Kappa, Mills College, 1969

Migration Patterns in Howling Monkeys, December, 2003

NSF grant #0414310 submitted 12/03, $658,000 over 5 years




Capture and behavior project on food selection and age in mantled howlers, February and June, 2001

supported by NSF Grant #9601766 to M. Teaford, Johns Hopkins Medical School, Baltimore MD




Observational methods comparison project, Hda La Pacifica, July-August, 1999

Assisted by D. Arden (Tulane). Capture and mark project with K.E. Glander, Duke University Primate Center




Behavioral research in conjunction with management of outdoor breeding colonies, July 1990 - June 1996

Develop research on physiology and behavior, continue collaboration on behavioral aspects of biomedical research. FDA Contract 223-88-1100, 223-91-1100, 223-94-1100 and SAIDS Supplemental Funding RR00164 Tulane Regional Primate Research Center




Behavioral research and management of FDA breeding contract and SAIDS breeding colony, July 1987 - June 1990

Associate Research Scientist, Delta Primate Center DAMD Grant 17-87-PP7863. Development of research into psychological well-being and environmental enrichment of nonhuman primates. Collaboration with other investigators on behavioral aspects of biomedical research. Taught “Human Origins” for University College, Tulane University




Research Fellow, Delta Primate Center, September 1985 - June 1987

Animal behaviorist of FDA Contract 223-85-1100 to R.H. Wolf. Analyze records, coordinate behavioral observations for 22 field cages of M. mulatta. Recommendations for long-range behavioral management of colony. Develop studies to analyze "psychological well-being" of caged primates.




Research Associate, Audubon Park and Zoological Garden, October 1984 - June 1985

Behavioral research on caged talapoins (Cercopithecus talapoin) and howlers (Alouatta caraya)




Research on heart rate reactivity and development of atherosclerosis in Macaca fascicularis, analyzing behavioral data on juvenile Erythrocebus patas and Cercopithecus aethiops, February, 1982 - January, 1983

NIH traineeship, Department of Pathology and Comparative Medicine, Bowman Gray School of Medicine, Winston-Salem, NC 27103. National Research Service Award #HLO7115.




Research project on female juvenile bonnet macaques (Macaca radiata), October, 1975 - September, 1976

Supported by USPHS Grant #RR00169. Research design and analysis. Trained six assistants in observation techniques and preliminary data analysis




Master's degree (library research) “Maintenance of squirrel monkey circannual cycles in natural, semi-natural, and laboratory environments.", December, 1970 - June, 1971

Involved with Chronobiology Laboratory (F. Halberg), University of Minnesota




Need for Achievement in Black Children Integrated Since Kindergarten Versus Black Children Integrated Since the Start of Third Grade, February, 1969 - June, 1969

Designed and carried out in Berkeley Public Schools. Utilized matched pairs of children for Aronson's Doodle Test




"Daily Behavior Patterns of Spider Monkeys (Ateles geoffroyi) at the San Francisco Zoo, January, 1969

January Term Project, Mills College




Undergraduate reader in Physical and Cultural Anthropology, September, 1968 - June, 1969

• Field Work, field research on howling monkeys in Costa Rica, September, 2002 - August, 2003


• Field Work, field research on “Female-Female Competition in Howling Monkeys, February, 2001 and June-2001-2002


• Undergraduate field research projects at Hda La Pacifica, June-July, 2000
  Supported by Newcomb Foundation, Kenneth J. Opat Fund and the Neotropical Ecology Institute of the Stone Center for Latin American Studies


• Complete census of howling monkey population of La Pacifica, Costa Rica, July-August, 1998
  Funded by National Geographic Research Grant 6248-98 and Faculty Summer Grant from the Roger Thayer Stone Center for Latin American Studies, Tulane University


• Behavioral observations as well as capturing, marking and bleeding howlers for behavioral, demographic, genetic, and endocrine studies, June/July/August 1985 - 1997
  Supported in part by Earthwatch and National Geographic. (Grant # 4546-91)


• Complete census of field site on howlers, June, 1984 - July, 1984
  Taught field course on Primate Social Behavior for School for Field Studies, Cambridge, MA. Four weeks in the field in Costa Rica. Also carried out complete census of field site on howlers; part of long-term project on Alouatta palliata


• Field research on howling monkeys (Alouatta palliata), November, 1978 - September, 1980
  Supported by NSF doctoral dissertation improvement grant #BNS78-19308, Explorers Club Grant, Graduate School Funds, and Departmental Funds. Daily behavioral observations, censusing, data tabulation, and preliminary analysis. All animals captured and marked at regular intervals during study. Advisor, PETER S. RODMAN, University of California, Davis. Field Supervisor, KENNETH E. GLANDER, Dept of Biological Anthropology and Anatomy, Duke University, Durham, North Carolina.

• American Society of Primatologists, Research and Development Committee Member, 2002 - present


• Tulane Primate Center IACUC, 1985 - 1996

  Departmental Committees/Service

• Stone Center Graduate Research Committee, Member, 2001 - 2002


• Committee for Andrew W. Mellon Visiting Scholar, Member, 1999 - 2000


• Executive Committee, Neotropical Ecology Institute (CLAS), Member, 1998 - 2000


• Student Awards Committee, Neotropical Ecology Institute (CLAS), Chair, 1997 - 2000


• Center for Women’s Studies Advisory Committee, Member, 1997 - 1998


• Primate Behavior and Ecology Program, Member, 2002 - 2003


• Tulane College, Freshman Advisor, 1997 - 1999

  University Committees/Service

• Newcomb College Honor Board, Member, 2000 - 2002


• Senate Committee on Collections, 1997 - 2002


• Graduate Teaching Evaluation (GSTOC), Member, 1998 - 2000

  Service to the Professional Community

  Journals/Volumes Refereed

• AMERICAN JOURNAL OF PRIMATOLOGY, Referee, 2004 - present

• ZOO BIOLOGY, Referee, 2004 - present

• FOLIA PRIMATOLOGICA, Referee, 2004 - present

• INTERNATIONAL JOURNAL OF COMPARATIVE PSYCHOLOGY, Referee, 2004 - present

• BRAIN BEHAVIOR AND IMMUNITY, Referee, 2004 - present

• LABORATORY ANIMAL SCIENCE, Referee, 2004 - present

• CONTEMPORARY TOPICS IN LABORATORY ANIMAL SCIENCE, Referee, 2004 - present

• CANADIAN JOURNAL OF ZOOLOGY, Referee, 2004 - present

• PSYCHOLOGICAL REVIEW, Referee, 2004 - present

• BEHAVIORAL ECOLOGY AND SOCIOBIOLOGY, Referee, 2004 - present

• PHYSIOLOGY AND BEHAVIOR, Referee, 2004 - present

• INTERNATIONAL JOURNAL OF PRIMATOLOGY, Referee, 2004 - present

• AMERICAN JOURNAL OF PRIMATOLOGY, Associate Editor, 1999 - 2004

• PHYSICAL ANTHROPOLOGY the official newsletter of the American Association of Physical Anthropologists, Editorial Advisory Board, 1999 - 2002

• AMERICAN JOURNAL OF PRIMATOLOGY, Consulting Editor, 1989 - 1999

• National Geographic Research, Proposal Reviewer, 2004 - present

• L.S.B. Leakey Foundation, Proposal Reviewer, 2004 - present

• National Science Foundation, Proposal Reviewer, 2004 - present

• Center for Field Studies (Earthwatch), Proposal Reviewer, 2004 - present

• Animal Behavior Society, Certified Applied Animal Behaviorist, 1988 - present

• American Society of Primatologists, Conservation Committee Member, 1998 - 2002

• American Association of Physical Anthropologists, Program Committee Member, 1998 - 2000

• Local Arrangements Committee, Annual Meeting of the American Society of Primatologists, Chair, August 12-16, 1999

• Elizabeth S. Watts Graduate Fellowship in Nonhuman Primate Growth and Development Committee, Amer. Soc.of Primatologists, Chair, 1994 - 1999

• American Society of Primatologists, Awards and Recognition Committee Member, 1990-1992, 1996-1998

• American Society of Primatologists, Executive Secretary, 1992 - 1996

• American Society of Primatologists, Board of Directors, 1992 - 1996

• American Society of Primatologists, Executive Committee Member, 1992 - 1996

• Audubon Zoo, Research Advisory Board, 1984 - 1995

Professional Affiliations

Professional Societies:
Member, American Association of Physical Anthropologists
Fellow, American Anthropological Association
Fellow, Human Biology Association
Animal Behavior Society
American Society of Primatologists
International Society of Primatology
International Society of Applied Ethology
Psychoneuroimmunology Research Society
Full member, Sigma Xi
Member, American Assoc. Adv. Science

Publications

Papers Published

1. Teaford, M.F., Clarke,M.R., and Glander, K.E., From the mouths of monkeys come tales of their lives', NATURAL HISTORY (2004).


2. Huynen MC, Clarke, MR, Social rank and interactions with males of adult female rhesus monkeys (Macaca mulatta) in a single-male versus a multi-male group (2004) (to book if accepted.).


3. Clarke, M.R., Crockett, C.M., Zucker, E.L., Zaldivar, M., A comparison of methods to survey free-ranging monkeys in the Costa Rican dry forest, LABORATORY PRIMATE NEWSLETTER, vol. 40 no. 4 (2001), pp. 4-6.


4. Clarke, M.R., Tremblay, A.M., and Arden, D.H., A comparison of methods to observe juvenile and group behavior in mantled howling monkeys, LABORATORY PRIMATE NEWSLETTER, vol. 39 no. 4 (2000), pp. 6-8.


5. Baskin, G.B., Ratterree, M., Davison, B.B., Falkenstein, K.P., Clarke, MR., England, J.D., Banier, M.T., Luzi, P., Rafi, M.A., Wenger, D.A., Genetic galactocerebrosidase deficiency (Globoid Cell Leukodystrophy, Krabbe Disease) in rhesus monkeys (Macaca mulatta), LABORATORY ANIMAL SCIENCE, vol. 48 no. 5 (1998), pp. 476-482.


6. Zucker, E.L., Clarke, M.R., Glander, K.E. and Scott, N.J., Jr., Sizes of home ranges and groups of mantled howling monkeys at La Pacifica: 1972-1991, BRENESIA (1996), pp. 45-46.


7. Ghoneim, G.M.; Shaaban, A.M.; Clarke, M.R., Irritable bladder syndrome in an animal model: A continuous monitoring study, NEUROUROLOGY AND URODYNAMICS, vol. 14 (1995), pp. 657-665.


8. Clarke MR & Cogswell FB., A comparison of two protocols to monitor parasitism in socially housed rhesus monkeys, CONTEMPORARY TOPICS IN LABORATORY ANIMAL SCIENCE, vol. 34 no. 5 (1995), pp. 82-83.


9. Clarke, M.R., Blanchard, J.L., Snyder, J.A., Infant-killing in pigtailed macaques: A colony management concern, LABORATORY PRIMATE NEWSLETTER, vol. 34 no. 4 (1995), pp. 1-3.


10. Clarke MR, Zucker EL and Glander KE, Group takeover by a natal male howling monkey (Alouatta palliata) and associated disappearance and injuries of immatures, PRIMATES, vol. 35 no. S4 (1994), pp. 435-442.


11. Clarke MR, Blanchard JL., All-male social group formation: Does cutting canine teeth promote social integration?, LABORATORY PRIMATE NEWSLETTER, vol. 33 no. 2 (1994), pp. 5-8.


12. Zucker E.L., Clarke M.R., Schilling P.W., Use of Natural and Provided Shade by Free-ranging Rhesus Monkeys on Raccoon Key, Florida: Management Considerations, CONTEMPORARY TOPICS IN LABORATORY ANIMAL SCIENCE, vol. 32 no. 3 (1993), pp. 6-8.


13. Zucker E.L. and Clarke M.R., Visitors’ movement patterns at conference poster sessions, vol. 76 (1993), pp. 212-214.


14. Zucker E.L. and Clarke M.R., Developmental and comparative aspects of social play of mantled howling monkeys in Costa Rica, BEHAVIOUR, vol. 123 no. 1-2 (1992), pp. 144-171.


15. Clarke M.R and Mayeaux D.J., Aggressive and affiliative behavior in green monkeys with differing housing complexity, AGGRESSIVE BEHAVIOR, vol. 18 (1992), pp. 231-239.


16. Phillippi-Falkenstein, K. and Clarke, M.R., A procedure for training corral-living rhesus monkeys for fecal and blood sample collection, LABORATORY ANIMAL SCIENCE, vol. 42 no. 1 (1992), pp. 83-85.


17. Clarke MR, Behavioral development and socialization of infants in a free-ranging group of howling monkeys (Alouatta palliata), FOLIA PRIMATOLOGICA, vol. 54 (1990), pp. 1-15.


18. Ratterree MS, Didier PJ, Blanchard JL, Clarke MR, Schaeffer D, Vitamin C deficiency in captive nonhuman primates fed commercial primate diet, LAB ANIMAL SCIENCE, vol. 40 no. 2 (1990), pp. 165-168.


19. Breeding colonies of macaques: A 1988 update, LABORATORY PRIMATE NEWSLETTER, edited by Clarke MR, vol. 28 no. 1 (1989), pp. 8-14.


20. Harrison RM, Clarke MR, Waggoner, R., Determining sperm quality in rhesus monkeys: Timing is all, LAB ANIMAL, vol. 18 no. 1 (1989), pp. 30-31.


21. Armstrong E, Clarke MR, and Hill EM, Relative size of the anterior thalamic nuclei differentiates anthropoids by social organization, BRAIN, BEHAVIOR & EVOLUTION, vol. 30 (1987), pp. 263-271.


22. Clarke MR, Kaplan JR, Nicks EC, and Grady AR, Use of space by young African greens (Cercopithecus aethiops spp.) and patas (Erythrocebus patas) monkeys in captivity, in COMPARATIVE BEHAVIOR OF AFRICAN MONKEYS, edited by E.L. Zucker (1987), pp. 81-97, Alan R. Liss, New York.


23. Clarke MR, and Glander KE, Female reproductive success in a group of free-ranging howling monkeys (Alouatta palliata) in Costa Rica, in FEMALE PRIMATES: STUDIES BY WOMEN PRIMATOLOGISTS, edited by M.F.Small (1984), pp. 111-126, Alan R. Liss, New York.


24. Clarke MR, Socialization, infant mortality, and infant-nonmother interactions in howling monkeys (Alouatta palliata) in Costa Rica, DISSERTATION ABSTRACTS INTERNATIONAL A43(4):1217, (Order #DA-8220132) (1982), University Microfilms, Inc., Ann Arbor, MI 48106.


25. Clarke MR, Social interactions of juvenile female bonnet monkeys (Macaca radiata), PRIMATES, vol. 19 no. 3 (1978), pp. 517-524.

Papers In Preparation

1. Clarke, M.R., Arden, D.H., Epstein, D.S., A comparison of activity patterns in adult male howling monkeys (Alouatta palliata) in three social groups in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY (2004) (In Press.).


2. Epstein, D.S. and Clarke, M.R., Proximity patterns in adult male howling monkeys, NEOTROPICAL PRIMATES (2004).


3. Clarke, M.R. and Glander, K.E., Juvenile emigration patterns in howling monkeys: Effect of presence of mother, ANIMAL BEHAVIOUR (2004).


4. Clarke, M.R. and Glander, K.E., Migration patterns in adult male and female howling monkeys: A 30 year study, BEHAVIOUR (2004).


5. Teaford, M.F., Clarke, M.R., Zilstra, M. and Glander, K.E., Analysis of Fecal and Stomach Contents of Howling Monkeys, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (2004).


6. Zucker EL and Clarke MR, Male-male interaction patterns in mantled howling monkeys, AMERICAN JOURNAL OF PRIMATOLOGY (2004).


7. Zucker, E.L., Clarke, M.R., Glander, K.E., Latencies to first births by immigrant female howling monkeys (Alouatta palliata) in Costa Rica (2004).


8. Clarke MR, Bertsch J, Phillippi, K., Carr DJJ, Changes in behavior and immune function following spontaneous rank reversal in rhesus monkeys (2004) (to book if accepted.).


9. Clarke MR, Interactions of adult female howlers with group infants in free-ranging howlers: Helping or harming? (2004).


10. Clarke MR, Adult male-infant interactions in a group of free-ranging howlers (2004).


11. Clarke MR, Parent offspring conflict in howlers (Alouatta palliata) (2004).


12. Clarke MR, Harrison RM, Zucker EL, Hormonal correlates of female behavior in free-ranging howling monkeys (2004).


13. Clarke MR and Harrison RM, Reproductive suppression in socially housed juvenile male rhesus monkeys (2004).


14. Clarke MR, Phillippi KM, and Zucker EL, . Male-male interaction patterns in captive Chinese male rhesus monkeys (2004).


15. Clarke MR & Daigle RM, A comparison of two strategies of social group reorganization in rhesus monkeys (2004).


16. Clarke MR, Kinship or familiarity? A comparison of social interactions in adult female rhesus monkeys (2004).


17. Clarke MR & Huynen MC, Social interactions of juvenile male rhesus monkeys and subsequent success in social group introduction (2004).


18. Clarke, M.R. and O’Neil, J.A., Behavioral comparison of foster and normal rhesus monkey infants. (2004).


19. Huynen, M. C. and Clarke, M.R., Female reproductive strategies: Competition for limited resource or preferred mate? (2004).

Book Reviews

1. Clarke MR, Review of Origins of Intelligence, JOURNAL OF HUMAN BIOLOGY, vol. 13 no. 3 (2001), pp. 425-426.


2. Clarke MR, Review of Demonic Males, AGGRESSIVE BEHAVIOR, edited by R. Wrangham and D. Peterson, vol. 25 no. 4 (1999), pp. 305-307.


3. Clarke MR, Review of Primate Conservation in the Tropical Rainforest, AMERICAN ANTHROPOLOGIST, edited by C. Marsh and R. Mittermeier, vol. 90 no. 1 (1988), pp. 171.

Journal Articles

1. Zucker, E.L., Clarke, M.R., Longitudinal assessment of immature-to-adult ratios in two groups of Costa Rican mantled howlers (Alouatta palliata), INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 24 no. 1 (2003), pp. 87-101.


2. Clarke, M.R., Crockett, C.M., Zucker, E.L., Zaldivar, M., Mantled howler population of Hacienda La Pacifica, Costa Rica from 1991 to 1998: Effects of deforestation, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 56 (2002), pp. 155-163.


3. Clarke, M.R., Collins, D.A., Zucker, E.L., Adaptations to deforestation in a free-ranging group of mantled howlers (Alouatta palliata) in Costa Rica, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 23 no. 2 (2002), pp. 365-381.


4. Zucker, E.L., Clarke, M.R., Glander, K.E., Body weights of adult female Alouatta palliata in Costa Rica increase with age, NEOTROPICAL PRIMATES, vol. 9 no. 3 (2001).


5. Zucker, E.L., Clarke, M.R., Glander, K.E., Body weights before and after first pregnancies of immigrant adult female howling monkeys (Alouatta palliata) in Costa Rica, NEOTROPICAL PRIMATES, vol. 9 no. 2 (2001), pp. 57-60.


6. Clarke, M.R., O’Neil, J.A.S., Morphometric comparison of Chinese-origin and Indian-derived rhesus monkeys (Macaca mulatta), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 47 no. 4 (1999), pp. 335-246.


7. Clarke, M.R., Glander, K.E., Zucker, E.L., Infant-nonmother interactions in mantled howling monkeys, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 19 (1998), pp. 451-472.


8. Stuart, M., Pendergast, V., Rumfelt, S., Pierberg, S. Greenspan, L. , Glander, K. and Clarke, M.R., Parasites of wild howlers (Alouatta spp.), INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 19 (1998), pp. 493-512.


9. Zucker, E. L., Clarke, M.R., Agonistic and affiliative relationships among adult female howling monkeys (Alouatta palliata) in Costa Rica over a four-year period, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 19 (1998), pp. 433-450.


10. Clarke M.R,; Harrison, R.M.; Didier, E.S., Behavioral, immunological and hormonal responses associated with social change in rhesus monkeys (Macaca mulatta), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 39 (1996), pp. 223-233.


11. Clarke MR and Zucker EL, Survey of the howling monkey population of La Pacifica: A seven-year follow-up, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 15 no. 1 (1994), pp. 29-41.


12. Phillippi, KM and Clarke, MR, Survey of parasites of rhesus monkeys housed in small social groups, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 27 (1992), pp. 293-302.


13. Stuart MD, Greenspan LL, Glander KE, Clarke MR, A coprological survey of parasites of wild mantled howling monkeys, Alouatta palliata, JOURNAL OF WILDLIFE DISEASES, vol. 26 no. 4 (1990), pp. 547-549.


14. Clarke MR, Koritnik DR, Martin LN, Baskin, GB, Cage enrichment, physiology and behavior in nursery-reared rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. Supplement 1 (1989), pp. 53-57.


15. Clarke MR, Kaplan JR, Bumsted PT, and Koritnik DK, Social dominance and serum testosterone concentration in dyads of Macaca fascicularis, JOURNAL OF MEDICAL PRIMATOLOGY, vol. 15 (1986), pp. 419-432.


16. Clarke MR, Zucker EL, and Scott NJ, Jr., Population trends of the mantled howler groups at La Pacifica, Guanacaste, Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 11 (1986), pp. 79-88.


17. Clarke MR, Infant-killing and infant disappearance following male takeover in a group of howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 5 no. 3 (1983), pp. 241-247.


18. Clarke MR, and Glander KE, Adoption of infant howling monkeys (Alouatta palliata), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 1 no. 4 (1981), pp. 469-472.

Papers Presented/Symposia/Abstracts

1. Clarke, M. R. and Glander, K.E., Adult migration patterns in the mantled howlers of La Pacifica, AMERICAN JOURNAL OF PRIMATOLOGY, Supplement (2004).


2. Clarke, M.R., Group size resurgence in mantled howlers (Alouatta palliata), AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. Supplement 38 (2004), pp. 75-76.


3. Clarke, M.R. , Glander, K.E., Males really do follow females after all: Thirty years data on male migration in howling monkeys at La Pacifica, Guanacaste Province, Costa Rica, The Animal Behavior Society, Boise Idaho (July 19-24, 2003).


4. Clarke MR, Managing monkeys from an evolutionary perspective, The Animal Behavior Society, Bloomington IN (July 13-17, 2002).


5. Nisbett R.A., Rodriguez L.L., Clarke M.R., Glander K.E., Teaford M.F., Jimenez Sanchez C., Preliminary report on the epidemiology of vesicular stomatitis virus in a free-ranging population of Costa Rican howling monkeys, AMERICAN JOURNAL OF MEDICINE AND HYGIENE, vol. 67 no. 2 (2002), pp. 272-273.


6. Clarke, M.R., Teaford, M.F., Preliminary study on hormone measurement in dried feces of free-ranging howling monkeys (Alouatta palliata) in Costa Rica, Suppl 1:67, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 57 (2002).


7. Zucker, E.L., Clarke, M.R., Glander, K.E., Habitat mediates relationship between age and interbirth intervals in Costa Rican mantled howling monkeys (Alouatta palliata), Suppl 1:33, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 57 (2002).


8. Seltzer, R.G., York, H.P., Clarke, M.R., A survery of vervet monkey (Chlorocebus [Cercopithecus] aethiops sabaeus) populations at five sites in Nevis, West Indies, Suppl 1:69-70, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 57 (2002).


9. Clarke, M.R., Glander, K.E., . Female immigration patterns in mantled howling monkeys (Alouatta palliata) at La Pacifica, GTE, Costa Rica, edited by Supplement, 34:54, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (2002).


10. Epstein, D.S., Clarke, M.R., Male-female partner preferences in mantled howling monkeys (Alouatta palliata) on Hda La Pacifica, Guanacaste Province, Costa Rica, The Animal Behavior Society, Corvallis OR (July 14-18, 2001).


11. Arden, D.H., Clarke, M.R., Social interactions between adult males and juvenile howling monkeys (Alouatta palliata) in Costa Rica, The Animal Behavior Society, Corvallis OR (July 14-18, 2001).


12. Clarke, M.R., Arden, D.H., Epstein, D.S., Gilbert, M.H., Activity patterns of adult male howling monkeys: Comparison by age, habitat and group composition, Supplement 1:28, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 53 (2001).


13. Clarke, M.R. and Glander, K.E., Presence of mother and juvenile dispersal in free-ranging howling monkeys (Alouatta palliata) in the tropical dry forest of Costa Rica, Supplement 32:50, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (2001).


14. Clarke, M.R., Anderson, L.S., Porter, J. and Martin, L.M., Behavior, hormones, and immune correlates of socially stable/unstable housing in juvenile rhesus monkeys (Macaca mulatta), SUPPLEMENT 30, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (2000), pp. 125-126.


15. Clarke, M.R., Tremblay, A.M., and Arden, D.H., Comparison of observational methods for juvenile and group behavior in mantled howling monkeys, Supplement 1, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 42 (2000), pp. 50-51.


16. Clarke, M. R., Crockett, C.M., Zucker, E.L., A comparison of methods used to census mantled howlers in the Costa Rican dry forest, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 49 no. 1 (1999), pp. 43-44.


17. Huynen, M.C., Clarke, M.R., Female familiarity and relatedness in captive groups of rhesus monkeys (Macaca mulatta), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 49 no. 1 (1999), pp. 65.


18. Huynen, M.C., Clarke, M.R., Female age class and competition for mates in social groups of rhesus monkeys (Macaca mulatta), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 49 no. 1 (1999), pp. 64-65.


19. Zucker, E.L., Clarke, M.R., Immature-to-adult ratios over time in two groups of Costa Rican mantled howling monkeys (Alouatta palliata) Annual variations and comparison with population parameters, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 49 no. 1 (1999), pp. 115-116.


20. Glasgow, M.E., Williams, L.E., Clarke, M.R., Environmental effects of female social behavior during pregnancy and infant development in captive Squirrel monkeys (Saimir bolivienses): Implications for management, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 49 no. 1 (1999), pp. 58.


21. Clarke, M.R., Zucker, E.L., Crockett, C.M. , Zaldivar, M., Population parameters of the howling monkeys of La Pacifica, Supplement 27, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (1999), pp. 108-109.


22. Pielet, H. A., Clarke, M.R., Social interactions of pigtailed and rhesus monkeys, Supplement 27, AMERICAN JOURNAL OF PRIMATOLOGY (1999), pp. 221-222.


23. Zucker, E.L., Clarke, M.R., Glander, K.E., Life histories of adult female howling monkeys in Costa Rica: Age, weight and status, Supplement 25, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (1998), pp. 239.


24. Clarke, M.R., Collins, D.A., Zucker, E.L., Changes in daily activity pattern and rates of social interactions in a free-ranging group of mantled howling monkeys (Alouatta palliata) in Costa Rica following partial deforestation of their home range, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 45 no. 2 (1998), pp. 174-175.


25. Zucker, E.L., Clarke, M.R., Glander, K.E., Latencies to first births for immigrating adult female howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 42 (1997), pp. 158.


26. Clarke, M.R., Harrison, R.M., O’Neil, J.A.S., Zucker, E.L., Reproductive hormones and migration patterns in free-ranging juvenile howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 42 (1997), pp. 100-101.


27. Lewis, R.J., Huynen, M.C., Clarke, M.R., The effects of varying social power on pigtailed monkey (Macaca nemestrina) and rhesus monkey (Macaca mulatta) social dynamics, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 42 (1997), pp. 128.


28. Clarke, M.R., Zucker E.L., Phillippi-Falkenstein, K., Activity and proximity patterns of free-ranging male and female juvenile howling monkeys (Alouatta palliata), Supplement 24, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (1997), pp. 93.


29. Clarke, M. R.; Zucker, E.L.; Phillippi Falkenstein, K., Activity and proximity patterns of juvenile howling monkeys with and without mothers in free-ranging social groups, 16th Congress of the International Primatological Society and the 19th Converence of the American Society of Primatologists, Madison WI (August 11-16, 1996).


30. Zucker, E.L.; Clarke, M.R.; O’Neil, J.A.S., Harrison, R.M., . Fecal testosterone values for free-ranging male mantled howling monkeys, 16th Congress of the International Primatological Society and the 19th Converence of the American Society of Primatologists, Madison WI (August 11-16, 1996).


31. Lewis, R.J.; Huynen , M.C.; Clarke, M.R., Social interactions and activity patterns of adult female pigtailed monkeys and rhesus monkeys, 16th Congress of the International Primatological Society and the 19th Converence of the American Society of Primatologists, Madison WI (August 11-16, 1996).


32. Huynen, M.C.; Clarke,M.R., Female rhesus monkeys’ social interactions vary with proportion of males, 16th Congress of the International Primatological Society and the 19th Converence of the American Society of Primatologists, Madison WI (August 11-16, 1996).


33. Clarke, M.R. and Snyder, J.A., Behavior of foster infants vs. natural infants in socially-housed rhesus monkeys, Southwestern Psychological Assocation Meeting, Houston TX (April 4-6, 1996).


34. Clarke , M. R.; Snyder, J.A., Morphological comparison of Indian-origin and Chinese-origin rhesus monkeys (Macaca mulatta), Supplement 22, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (1996), pp. 86.


35. Clarke MR, Daigle R, Snyder J., A comparison of social interactions with kin, subgroup members and other members in two social groups of rhesus monkeys (Macaca mulatta), Southwestern Psychology Association Meeting, San Antonio, TX (April 12-15, 1995).


36. Huynen, M.C.; Clarke, M.R., Social interactions of high and low ranking adult female rhesus monkeys (Macaca mulatta) in one-male and multi-male social groups, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 32 no. 6 (1995), pp. 128.


37. Zucker EL, Clarke MR & Harrison RM., Fecal estradiol values for group-living cycling, pregnant and lactating female howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 36 no. 2 (1995), pp. 167.


38. Clarke M, Carr D & Porter J., A preliminary study on behavioral and immunological correlates of social instability in juvenile rhesus monkeys, Meeting of the Psychoneuroimmunology Research Society, Key Biscayne, FL (November 13-20, 1994).


39. Aertker, MW, Clarke, MR, and Ghoneim, GM, The effect of environmental change on micturition and behavior of rhesus monkeys, Society for Basic Urological Research, San Francisco CA (May 13-14, 1994).


40. Zucker EL & Clarke MR, Time budgets and proximity patterns of juvenile howling monkeys (Alouatta palliata) in Costa Rica, 31st Annual Meeting of the Animal Behavior Society, Seattle, WA (July 23-28, 1994).


41. Huynen MC & Clarke MR, Social interactions of high/low ranking adult female rhesus monkeys, edited by 31st Annual Meeting of the Animal Behavior Society, Seattle, WA (July 23-28, 1994).


42. Clarke MR, Daigle RM., Comparing success of two social reorganization strategies: When theory and management coincide, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 33 no. 3 (1994), pp. 201-202.


43. Zucker EL, Clarke MR, Putnam PM, Harrison RM, Validity of measures assessing reproductive status of female howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 33 no. 3 (1994), pp. 255.


44. Clarke MR, Bertsch J, Phillippi-Falkenstein K, and Carr DJJ, Social instability and immune function in female rhesus monkeys (Macaca mulatta) following social reorganization, Research Perspectives in Psychoneuroimmunology IV., Boulder, CO (April 21-24, 1993).


45. Clarke MR, Bertsch J, Phillippi-Falkenstein K and Carr DJJ, Social instability and natural killer cell function in female rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 30 no. 4 (1993), pp. 304.


46. Huynen MC and Clarke MR., Social interactions of young adult males housed in an all-male group, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 30 no. 4 (1993), pp. 319-320.


47. Zucker EL and Clarke MR, Social relationships of adult female howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 30 no. 4 (1993), pp. 361-2.


48. Clarke MR, Daigle RM, McGrew WC, Stone-handling and cheekpouch filling in a colony of rhesus monkeys, Supp 16, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY (1993), pp. 71.


49. Clarke MR, Harrison RM, Martin LN, Rearing history and survival time in SIV-infected juvenile rhesus monkeys, Nonhuman Primate Models for AIDS, San Juan, PR (November 1992).


50. Zucker, E.L., Clarke, M.R., Glander, KE, and Scott, N.J., Jr., Locations of mantled howling monkey groups at La Pacific, Costa Rica, 1972-1991, Animal Behavior Meetings, Kingston, Ontario (June, 1992).


51. Phillippi-Falkenstein K, Clarke MR, and Zucker EL., Adult Chinese male rhesus monkeys social interactions: a comparison of two social introduction protocols, Southwestern Psychological Association, Austin, TX (April 15-18, 1992).


52. Phillippi-Falkenstein K, Clarke MR and Cogswell FB, A comparison of two protocols to monitor parasitism in socially housed rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 27 (1992), pp. 51.


53. Clarke MR and Zucker EL, A new survey of the howling monkey population of La Pacifica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 27 (1992), pp. 23.


54. Clarke, M.R., Harrison, RM, and Didier, EJ., Behavioral and immunological correlates of social reorganization, . American Society of Primatoglogists, Veracruz, Mexico (June, 1991).


55. Clarke MR, Behavioral studies at the Tulane Regional Primate Research Center, Southeastern Psychological Association, New Orleans, LA (March 20-23, 1991).


56. Clarke MR, The effects of environmental complexity: Three Studies, Southwestern Psychological Association, New Orleans, LA (April 11-13, 1991).


57. Clarke MR, Zucker EL and Harrison RM, Fecal estradiol, sexual swelling, and sociosexual behavior of free-ranging female howling monkeys in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 24 no. 2 (1991), pp. 93.


58. Zucker EL, Clarke MR and Schilling PW, Use of natural and provided shade by free-ranging rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 24 no. 2 (1991), pp. 143-144.


59. Clarke MR and Harrison RM, Timing of maturation in male rhesus monkeys: effects of social environment, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 20 no. 3 (1990), pp. 180.


60. Clarke MR, Phillippi KM, Falkenstein JA, Moran EA, and Suomi SJ, Training corral-living rhesus monkeys for fecal and blood sample collection, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 20 no. 3 (1990), pp. 181.


61. Phillippi KM, Clarke MR, Blanchard JL, Survey of pathogenic and nonpathogenic parasites of rhesus monkeys housed in small social groups, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 20 no. 3 (1990), pp. 221.


62. Zucker EL, Mayeaux DJ, Phillippi KM and Clarke MR, Interactions of male Chinese rhesus monkeys while in an all-male group and in breeding groups, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 20 no. 3 (1990), pp. 248.


63. Mayeaux, D.J. and Clarke, MR, Effects of changing habitat complexity on social group formation in green monkeys, Southeastern Psychological Association, Houston TX (April, 1989).


64. Clarke MR, and Zucker EL, Social correlates of timing of sexual maturity in free-ranging howling monkeys (Alouatta palliata), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 18 no. 2 (1989), pp. 140.


65. Clarke MR, Martin LN, and Baskin GB, Cage enrichment, behavior, and physiology in nursery-reared rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 14 no. 4 (1988), pp. 416.


66. Clarke MR, Breeding colonies of macaques: Progress and recent developments, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 14 no. 4 (1988), pp. 415.


67. Stuart M, Bartlett L, Glander K, and Clarke M, Helminth parasites of mantled howler monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 14 no. 4 (1988), pp. 447.


68. Berg MA, and Clarke MR, Social behavior in two groups of rhesus monkeys, Southwestern Psychological Association, New Orleans, LA. (April, 1987).


69. Clarke MR, Social interactions of howling monkey infants with group members: A cross-species review, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 8 no. 5 (1987), pp. 427.


70. Clarke MR, and Baskin GB, Patterns of infant mortality in a rhesus (Macaca mulatta) breeding colony, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 12 no. 3 (1987), pp. 336-337.


71. Harrison RM, Clarke MR, and Waggoner R, Effect of sexual activity of response to electroejaculation in rhesus monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 12 no. 3 (1987), pp. 346-347.


72. Zucker EL, and Clarke MR, Patterns of male-female spatial relationships in a multi-male group of howling monkeys (Alouatta palliata) in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 12 no. 3 (1987), pp. 379-380.


73. Clarke MR, Effects of habitat enrichment on sociospatial behavior in a group of talapoin monkeys, In: Zoological Parks: Applications to Husbandry, Management, and Habitat Design. 32nd Annual meeting of the Southeastern Psychological Association, Orlando, Fl (March 26-29, 1986).


74. Clarke MR, and Watson EA, Jr., Reproductive success of feral vs. colony born rhesus females: Implications for breeding programs, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 10 (1986), pp. 443.


75. Zucker EL, Watson EA, Jr., and Clarke MR, Group change and secondary sex ratios for corral-living rhesus macaques (Macaca mulatta), AMERICAN JOURNAL OF PRIMATOLOGY, vol. 10 (1986), pp. 443.


76. Zucker EL, and Clarke MR, Male-male interactions in a group of free-ranging howling monkeys, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 10 (1986), pp. 443.


77. Clarke MR, Interactions of adult male howling monkeys (Alouatta palliata) with immatures in a free-ranging social group, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 69 (1986), pp. 188.


78. Clarke MR, Teaching primate behavior in a field situation, In: Beyond the Laboratory, Alternatives in Comparative Psychology. 31st Annual Meeting of the Southeastern Psychological Association, Atlanta, GA (March 27-30, 1985).


79. Clarke MR, Behavior of adult females towards group infants in free-ranging howlers, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 8 no. 4 (1985), pp. 336.


80. Clarke MR, and Zucker EL, Distribution of howling monkeys (Alouattapalliata) in a dry, deciduous tropical forest, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 66 no. 2 (1985), pp. 156.


81. Clarke MR, A comparison of male and female infant howling monkeys in a free-ranging group in Costa Rica, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 6 no. 4 (1984), pp. 401.


82. Clarke MR, and Glander KE, Variability in reproductive success of female mantled howlers over an 11-year period, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 4 no. 4 (1983), pp. 440.


83. Kaplan JR, Manuck SB, Clarke MR, and Bumsted PT, Psychophysiology of spontaneous social behavior among cynomolgus macaques, AMERICAN JOURNAL OF PRIMATOLOGY, vol. 4 no. 4 (1983), pp. 347.


84. Clarke MR, Nicks EC, Grady AR, and Kaplan JR, Use of space by juvenile green monkeys and patas monkeys. In: Comparative behavior of African monkeys: Are primate models applicable?, INTERNATIONAL JOURNAL OF PRIMATOLOGY, vol. 3 (1982), pp. 153.


85. Clarke MR, Aspects of male behavior in the mantled howler (Alouatta palliata Gray) in Costa Rica, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 54 (1981), pp. 209.


86. Glander KE, and Clarke MR, Sex ratio and differential mortality in howling monkeys, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 54 (1981), pp. 225.


87. Clarke MR, Interactions of bonnet monkey infants with nonmothers, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 50 (1979), pp. 428.


88. Clarke MR, Various Presentations Given (1977-2004) (Also at 1977, 1979, 1981, 1985, 1986, 1987. 1993. 1996 through 2004 Annual meetings of American Association Physical Anthropologists; 1982, 1988, and 1996 meeting of the International Society of Primatology; 1983-2002, and 2004 meetings of the American Society of Primatologists..).


89. Clarke MR, and Neville MK, Social interactions of female juvenile bonnet macaques, AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, vol. 47 (1977), pp. 124.