Richard B. Forward, Professor Emeritus of Zoology  

Richard B. Forward

R. Forward works on the physiological ecology of marine crustaceans and fishes. Studies focus on sensory physiology and behavioral responses to environmental (e.g. light, temperature, salinity) and chemical cues, and biological rhythms. Recent studies have applied result to diel vertical migration and selective tidal stream transport.

Ph.D., University of California at Santa Barbara, 1969
B.A., Stanford University, 1965
BS, Stanford University

Office Location: 135 Duke Marine Lab Rd, Beaufort, NC 28516
Office Phone: (252) 504-7610
Email Address:

marine science

Representative Publications   (More Publications)   (search)

  1. Cohen, JH; Cronin, TW; Lessios, N; Forward, RB, Visual physiology underlying orientation and diel behavior in the sand beach amphipod Talorchestia longicornis., The Journal of experimental biology, vol. 213 no. Pt 22 (November, 2010), pp. 3843-3851 [21037063], [doi]  [abs].
  2. Forward, R.B., Jr. and J. H. Cohen, Vertical migration of aquatic animals., in Encyclopedia of Animal Behavior. Breed, M. D. and Moore, J., eds. Academic Press, Oxford, vol. 3 (2010), pp. 485-490 .
  3. Forward, RB; Freeman, E; Gall, K; Saba, K, Seasonal changes in entrainment cues for the circadian rhythm of the supratidal amphipod Talorchestia longicornis., Marine and Freshwater Behaviour and Physiology, vol. 43 no. 305-319 (2010), pp. 305-319 [doi]  [abs].
  4. Walsh, ME; Bourla, MH; Sabella, CM; Forward Jr, RB, Y-axis orientation in the supratidal amphipod Talorchestia longicornis (Say): Hierarchy of sun, beach slope and landmarks as orientation cues., Marine and Freshwater Behavior and Physiology, vol. 43 (2010), pp. 203-220 .

Curriculum Vitae